Trevor’s publications

Peer-reviewed papers

150. Worthy, T. H. 2012. A phabine pigeon (Aves: Columbidae) from Oligo-Miocene Australia. Emu 112: 23-31. ERA10B.

149. Worthy, T. H. 2012. A new species of Oligo-Miocene darter (Aves: Anhingidae) from Australia. The Auk 129: 96-104. ERA10A.

148. Worthy, T. H., and R. P. Scofield. In press January 2012. 21st century advances in knowledge of the biology of moa (Aves: Dinornithiformes), Diagnoses revised, and a new morphological analysis. New Zealand Journal of Zoology *****. ERA10C.

147. Schwarzhans, W., R.P. Scofield, A.J.D. Tennyson, J.P. Worthy, and T.H. Worthy, in press 22 March 2011. Fish remains, mostly otoliths, from the non-marine Early Miocene of Otago, New Zealand. Acta Palaeontologica Polonica ***********. ERA10B.

146. Gillespie, R., A. B. Camens, T. H. Worthy, N. J. Rawlence, C. Reid, F. Bertuch, V. Levchenko and A. Cooper, in press Jan 2012. Man and megafauna in Tasmania: closing the gap. Quaternary Science Reviews *** ERA10A*.

145. Fitzgerald, E. M. G., T. Park, T. H. Worthy, in press. First giant bony-toothed bird (Pelagornithidae) from Australia. Journal of Vertebrate Paleontology. Accepted 3 February 2012. ERA10A.

144. Wood, J. R., J. M. Wilmshurst, T. H. Worthy, A. Cooper, in press. First coprolite evidence for the diet of Anomalopteryx didiformis, an extinct forest ratite from New Zealand. New Zealand Journal of Ecology 36: ***********, accepted 8 February 2012. ERA10B.

143. Lee, M. S. Y. and T. H. Worthy. 2012. Likelihood reinstates Archaeopteryx as a primitive bird. Biology Letters 8(2): 308-311; published ahead of print September 21, 2011; doi:10.1098/rsbl.2011.0760. ERA10A. (Featured in Nature Research Highlights 479:8, 3 Nov. 2011, doi:10.1038/479008a ;  http://www.nature.com/nature/journal/v479/n7371/full/479008a.html)
142. Irwin, G., T.H. Worthy, S. Best, S. Hawkins, J. Carpenter & S. Matararaba 2011. Further investigations at the Naigani Lapita site (VL 21/5), Fiji: Excavation, radiocarbon dating and palaeofaunal extinction. Journal of Pacific Archaeology 2(2): 66-78. ERA10B.

141. Worthy, T. H., A. J. D. Tennyson, and R. P. Scofield, 2011. An Early Miocene diversity of parrots (Aves, Strigopidae, Nestorinae) from New Zealand. Journal of Vertebrate Paleontology 31: 1102-1116. ERA10A.

140. Worthy, T.H., 2011. Descriptions and phylogenetic relationships of a new genus and two new species of Oligo-Miocene cormorants (Aves: Phalacrocoracidae) from Australia. Zoological Journal of the Linnean Society 163: 277-314. ERA10A.

139. Rawlence, N. J., R. P. Scofield, J. R. Wood, J. M. Wilmshurst, N. T. Moar, T. H. Worthy 2011. New palaeontological data from the excavation of the Late Glacial Glencrieff miring bone deposit, North Canterbury, South Island, New Zealand. Journal of the Royal Society of New Zealand 41: 217-236. ERA10C.

138. Worthy, T.H., W. E. Boles. 2011. Australlus, a new genus for Gallinula disneyi (Aves: Rallidae) and a description of a new species from Oligo-Miocene deposits at Riversleigh, Northwestern Queensland, Australia. Records of the Australian Museum 63(1): 61-77. ERA10C. DOI:10.3853/j.0067-1975.63.2011.1563

137. Worthy, T.H., A.J.D. Tennyson, R.P. Scofield, 2011. Fossils reveal an early Miocene presence of the aberrant gruiform Aves: Aptornithidae in New Zealand. Journal of Ornithology 152: 669-680, DOI: 10.1007/s10336-011-0649-6. ERA10B.

136. Wood J. R., J. M. Wilmshurst, T. H. Worthy, A. Cooper, 2011. Sporormiella as a proxy for non-mammalian herbivores in island ecosystems. Quaternary Science Reviews 30: 915-920. ERA10A*.

135. Worthy T.H., A.J.D. Tennyson, S.J. Hand, H. Godthelp, R.P. Scofield 2011. Terrestrial turtle fossils from New Zealand refloat Moa’s Ark. Copeia 2011(1): 72-76. ERA10B.

134. Worthy, T.H. & R. Bollt, 2011. Prehistoric Birds and Bats from the Atiahara Site, Tubuai, Austral Islands, East Polynesia. Pacific Science 65(1): 69-85. doi: 10.2984/65.1.069 ERA10C.

133. White, A.W., T.H. Worthy, S. Hawkins, S. Bedford, & M. Spriggs. 2010. Megafaunal meiolaniid horned turtles survived until early human settlement in Vanuatu, Southwest Pacific. Proceedings of the National Academy of Sciences of the United States of America 107(350): 15512-15516. ERA10A*. www.pnas.org/cgi/doi/10.1073/pnas.1005780107

132. Hyde, N.H.S.; T.H. Worthy. 2010. The diet of New Zealand falcons (Falco novaeseelandiae) on the Auckland Islands, New Zealand. Notornis 57: 19-26. ERA10B.

131. Tennyson, A.J.D., R.L. Palma, R.P. Scofield, T.H. Worthy, 2010. Emending the species name for Lopdells’ penguin. Notornis 57: 54-55. ERA10B.

130. Checklist Committee, Ornithological Society of New Zealand; B.J. Gill (convener), B.D. Bell, G.K. Chambers, D.G. Medway, R.P. Scofield, A.J.D. Tennyson, T.H. Worthy. 2010. Letter supporting Case 3499 ‘Comments on the proposed conservation of Anthochaera Vigors & Horsfield, 1827 and Philesturnus Geoffroy Saint-Hilaire, 1832 (Aves) by suppression of the generic name Creadion Vieillot, 1816′. Bulletin of Zoological Nomenclature 67: 94. ERA10C.

129. Holland, B.R., H.G. Spencer, T.H. Worthy & M. Kennedy, 2010. Identifying Cliques of Convergent Characters: Concerted Evolution in the Cormorants and Shags. Systematic Biology 59:1-13. ERA10A* DOI:10.1093/sysbio/syq023.

128. Worthy, T.H., S.J. Hand, J.M.T. Nguyen, A.J.D. Tennyson, J.P. Worthy, R.P. Scofield W.E. Boles & M. Archer, 2010. Biogeographical and phylogenetic implications of an Early Miocene wren (Aves: Passeriformes: Acanthisittidae) from New Zealand. Journal of Vertebrate Paleontology 30: 479-498; DOI: 10.1080/02724631003618033. ERA10A.

127. Anderson, A., C. Sand, F. Petchey, & T.H. Worthy, 2010. Faunal extinction and human habitation in New Caledonia: initial results and implications of new research at the Pindai Caves. Journal of Pacific Archaeology 1:89-109. ERA10B.

126. Shepherd, L.D., J. Cooper, J. Haile, P. Scofield, A.J.D. Tennyson, T.H. Worthy, 2009. Selection of a neotype for Apteryx mantelli Bartlett, 1852, with the support of genetic data. Bulletin of the British Ornithologist’s Club 129(4): 195-197.

125. Worthy, T.H., 2009. Descriptions and phylogenetic relationships of two new genera and four new species of Oligo-Miocene waterfowl (Aves: Anatidae) from Australia. Zoological Journal of the Linnean Society 156: 411-454. ERA10A.

124. Bunce, M., T.H. Worthy, M.J. Phillips, R.N. Holdaway, E. Willerslev, J. Haile, B. Shapiro, R.P. Scofield, A. Drummond, P.J.J. Kamp & A. Cooper, 2009. The evolutionary history of the extinct ratite moa and New Zealand Neogene paleogeography. Proceeding of the National Academy of Sciences U. S. A. 106: 20646-20651. doi:10.1073/pnas.0906660106 ERA10A*.

123. Boessenkool, S., J.J. Austin, T.H. Worthy, P. Scofield, A. Cooper, P. J. Seddon & J.M. Waters, 2009. Relict or colonizer? Extinction and range expansion of penguins in southern New Zealand. Proceedings of the Royal Society B 276: 815-821. ERA10A*. doi:10.1098/rsbp.2008.1246 [online 20 Nov 2008; 2009 for nomenclature]

122. Jones, M.E.H., A.J.D. Tennyson, J.P. Worthy, S.E. Evans & T.H. Worthy, 2009. A sphenodontine (Rhynchocephalia) from the Miocene of New Zealand and palaeobiogeography of the tuatara (Sphenodon). Proceedings of the Royal Society B 276: 1385-1390 ERA10A*. (doi: 10.1098/rspb.2008.1785)

121. Worthy, T.H. & J. Scanlon, 2009. An Oligo-Miocene Magpie Goose Aves: Anseranatidae from Riversleigh, north western Queensland, Australia. Journal of Vertebrate Paleontology 29: 205-211. ERA10A.

120. Worthy, T.H., S.J. Hand, J.P. Worthy, A.J.D. Tennyson & R.P. Scofield 2009. A large fruit pigeon (Columbidae) from the Early Miocene of New Zealand. The Auk 126(3): 649-656. ERA10A.

119. Hand, S.J., V. Weisbecker, R.M.D. Beck, M. Archer, H. Godthelp, A.J.D. Tennyson & T.H. Worthy, 2009. Bats that walk: a new evolutionary hypothesis for the terrestrial behaviour of New Zealand’s endemic mystacinids. BMC Evolutionary Biology 2009, 9: 169. ERA10A. doi:10.1186/1471-2148-9-169.

118. Lee, M.S.Y., M.N. Hutchinson, T.H. Worthy, M. Archer, A.J.D. Tennyson, J.P. Worthy & R.P. Scofield, 2009. Miocene skinks and geckos reveal long-term conservatism of New Zealand’s lizard fauna. Biology Letters 5(6): 833-837, doi:10.1098/rsbl.2009.0440. ERA10A.

117. Worthy, T.H. & M.S.Y. Lee, 2008. Affinities of Miocene waterfowl (Anatidae: Manuherikia, Dunstanetta and Miotadorna) from the St Bathans Fauna, New Zealand. Palaeontology 51(3): 677-708. doi: 10.1111/j.1475-4983.2008.00778.x. ERA12.

116. Worthy, T.H., A.J.D. Tennyson, S.J. Hand & R.P. Scofield, 2008. A new species of the diving duck Manuherikia and evidence for geese (Aves: Anatidae: Anserinae) in the St Bathans Fauna (Early Miocene), New Zealand. Journal of the Royal Society of New Zealand 38(2): 97-114. ERA10C.

115. Wilmshurst, J.M., A.J. Anderson, T.F.G. Higham & T.H. Worthy, 2008. Dating the late prehistoric dispersal of Polynesians to New Zealand using the commensal Pacific rat. Proceeding of the National Academy of Sciences 105(22): 7676-7680. ERA10A*.

114. Wood, J.R., T.H. Worthy, N.J. Rawlence, S.M. Jones, R.N. Holdaway & S.E. Read, 2008. A deposition mechanism for Holocene miring bone deposits, South Island, New Zealand. Journal of Taphonomy 6: 1-20. ERA10C.

113. Worthy, T.H., 2008. Pliocene waterfowl (Aves: Anseriformes) from South Australia and a new genus and species. Emu 108: 153-165. ERA10B.

112. Wood, J.R., N.J. Rawlence, G. Rogers, J. Austin, A. Cooper & T.H., Worthy, 2008. Coprolite deposits reveal the diet and ecology of the extinct New Zealand megaherbivore moas (Aves, Dinornithiformes). Quaternary Science Reviews 27: 2593-2602. ERA10A*.

111. Worthy, T.H. S.L. Olson, & T. Smith, 2008. A reassessment of the fossil goose Anser scaldii Lambrecht, 1933 (Aves: Anatidae). Bulletin of the British Ornithologist’s Club 128(4): 228-232.

110. Worthy, T.H., A.J.D. Tennyson, C. Jones, J.A. McNamara, & B.J. Douglas, 2007. Miocene waterfowl and other birds from Central Otago, New Zealand. Journal of Systematic Palaeontology 5(1): 1-39. ERA10B. On line 14 Nov 2006; doi:1017/S1477201906001957.

109. Nunn, P.D., T. Ishimura, W.R. Dickinson, K. Katayama, F. Thomas, R. Kumar, S. Matararaba, J. Davidson & T. Worthy, 2007. The Lapita occupation at Naitabale, Moturiki Island, central Fiji. Asian Perspectives 46: 96-132. ERA10B.

108. Worthy, T.H. & N.S. Pledge, 2007. A shelduck (Anatidae: Tadorna) from the Pliocene of South Australia. Transactions of the Royal Society of South Australia 131: 107-115. ERA10C.

107. Worthy, T.H., 2007. Letters. Moas and phylogenomics: How nomenclatural errors do a disservice to the understanding of moa taxonomy. Auk 124: 1447-1449. ERA10A.

106. Worthy, T.H., 2007. Extinction & Biogeography of Tropical Pacific Birds. Invited review. Emu 107: 248-251. ERA10B.

105. Worthy, T.H., A.J.D. Tennyson, M. Archer, A.M. Musser, S.J. Hand, C. Jones, B.J. Douglas, J.A. McNamara & R.M.D. Beck. 2006. Miocene mammal reveals a Mesozoic ghost lineage on insular New Zealand, southwest Pacific. Proceeding of the National Academy of Sciences 103(51): 19419-19423.

104. Wragg, G.M. & T.H. Worthy, 2006. A new species of extinct imperial pigeon (Ducula: Columbidae) from Henderson Island, Pitcairn Group. Historical Biology 18(2): 127-140.

103. Worthy, T.H., 2005. A new species of Oxyura (Aves: Anatidae) from the New Zealand Holocene. Memoirs of the Queensland Museum 51: 255-272.

102. Worthy, T.H., 2005. Rediscovery of the types of Dinornis curtus Owen and Palapteryx geranoides Owen, with a new synonymy (Aves: Dinornithiformes) Tuhinga: Records of the Museum of New Zealand 16: 57-67.

101. Scofield, R. P., McAllan, I. A. W., Palma, R., Worthy, T.H. 2005. Stabilisation of the genus-group name of the New Zealand bellbird. Notornis 52: 171-172.

100. Worthy, T.H. & P. Scofield, 2004. Skeletal and dental variation within and between Mystacina species in southern New Zealand. New Zealand Journal of Zoology 31: 351-361.

99. Anderson, A., P. Scofield & T. Worthy, 2004. The number of moa at Wairau Bar: Correction and comment. Records of the Canterbury Museum 18: 49-50.

98. Worthy, T.H., 2004. Letter to the editor. Journal of the Royal Society of New Zealand 34: 105-106.

97. Worthy, T.H., 2004. The Holocene fossil waterfowl fauna of Lake Poukawa, North Island, New Zealand. Tuhinga: Records of the Museum of New Zealand 15: 77-120.

96. Worthy, T.H., 2004. The fossil rails (Aves: Rallidae) of Fiji with descriptions of a new genus and species. Journal of the Royal Society of New Zealand 34(3): 295-314.

95. Scofield, P., T.H. Worthy & H. Schlumpf, 2003. What birds were New Zealand’s first people eating? Wairau Bar’s avian remains re-examined. Records of the Canterbury Museum 17: 17-35.

94. Pregill G.K. & T.H. Worthy, 2003. A new iguanid lizard (Squamata, Iguanidae) from the late Quaternary of Fiji, Southwest Pacific. Herpetologica 59: 57-67.

93. Worthy, T.H. & J.A. Grant-Mackie, 2003. Late Pleistocene avifaunas from Cape Wanbrow, Otago, South Island, New Zealand. Journal of the Royal Society of New Zealand 33: 427-485.

92. Worthy, T.H., 2003. A new extinct species of snipe Coenocorypha from Vitilevu, Fiji. Bulletin of the British Ornithologists Club 123(2): 90-103.

91. Palma, R.L., T.H. Worthy & A.J.D. Tennyson, 2003. Resolution of the status of the taxon Apteryx maxima. Tuhinga: Records of the Museum of New Zealand 14: 1-9.

90. Worthy, T.H. & D. Roscoe, 2003. Takaka Fossil Cave – a stratified Late Glacial to Late Holocene deposit from Takaka Hill, New Zealand. Tuhinga: Records of the Museum of New Zealand 14: 41-60.

89. Bunce, M., T.H. Worthy, T. Ford, W. Hoppitt, E. Willerslev, A. Drummond, & A. Cooper, 2003. Extreme reversed sexual size dimorphism in the extinct New Zealand moa Dinornis. Nature 425: 172-175.

88. Worthy, T.H. & G.M. Wragg, 2003. A new species of Gallicolumba: Columbidae from Henderson Island, Pitcairn Group. Journal of the Royal Society of New Zealand 33: 769-793.

87. Tennyson, A.J.D., R.L. Palma, H. Robertson, T.H. Worthy, & B. J. Gill, 2003. A new species of kiwi (Aves, Apterygiformes) from Okarito, New Zealand. Records of the Auckland Museum 40: 55-64.

86. Worthy T.H. & B.J. Gill, 2002. New distributional records of the extinct New Zealand duck Malacorhynchus scarletti (Anatidae). Records of the Auckland Museum 39: 49-52.

85. Worthy, T.H., R. Walter, & A.J. Anderson, 2002. Fossil fauna from Niue Island. Appendix 2, pp 165-167. In, R. Walter & A. Anderson (eds), The archaeology of Niue Island West Polynesia. Bishop Museum Bulletins in Anthropology 10, x+167 pp.

84. Worthy, T.H.; Swabey, S. E. J. 2002. Avifaunal changes revealed in Quaternary deposits near Waitomo Caves, North Island, New Zealand. Journal of the Royal Society of New Zealand 32: 293-325.

83. Duncan, R.P., T.M. Blackburn, T.H. Worthy, 2002. Prehistoric bird extinctions and human hunting. Proceedings of the Royal Society London Series B. 269: 517-521.

82. Worthy, T.H. & S.L. Olson, 2002. Relationships, adaptations, and habits of the extinct duck ‘Euryanas’ finschi. Notornis 49: 1-17.

81. Worthy, T.H., 2002. The New Zealand musk duck (Biziura delautouri Forbes 1892). Notornis 49: 19-28.

80. Worthy, T.H., 2002. The youngest giant: discovery and significance of the remains of a giant moa (Dinornis giganteus) near Turangi, in central North Island, New Zealand. Journal of the Royal Society of New Zealand 32: 183-187.

79. Worthy, T.H., C.M. Miskelly, & (Bob) R.A. Ching, 2002. Taxonomy of North and South Island snipe (Aves: Scolopacidae: Coenocorypha), with analysis of a remarkable collection of snipe bones from Greymouth, New Zealand. New Zealand Journal of Zoology 29: 231-244.

78. Worthy, T.H., R.N. Holdaway, B.V. Alloway, J. Jones, J. Winn, & D. Turner, 2002. A rich Pleistocene – Holocene avifaunal sequence from Te Waka #1: terrestrial fossil vertebrate faunas from inland Hawke’s Bay, North Island, New Zealand. Part 2. Tuhinga, Records of the Museum of New Zealand Te Papa Tongarewa 13: 1-38.

77. Molnar, R. E.; Worthy, T.H.; Willis, P. M. A. 2002. An extinct Pleistocene endemic mekosuchine crocodylian from Fiji. Journal of Vertebrate Paleontology 22(3): 612-628.

76. Holdaway, R.N., R.G. Roberts, N.R. Beavan-Athfield, J.M. Olley & T.H. Worthy, 2002. Optical dating of quartz sediments and accelerator mass spectrometry 14C dating of bone gelatin and moa eggshell: a comparison of age estimates for non-archaeological deposits in New Zealand. Journal of the Royal Society of New Zealand 32(3): 463-505.

75. Worthy, T.H., 2001. A giant flightless pigeon gen. et sp. nov. and a new species of Ducula (Aves: Columbidae), from Quaternary deposits in Fiji. Journal of the Royal Society of New Zealand 31: 763-794.

74. Worthy, T.H. 2001. Quaternary fossil faunas of Otago, South Island, New Zealand. Errata. Journal of the Royal Society of New Zealand 31(3): 685-686.

73. Worthy, T.H., 2001. A fossil vertebrate fauna accumulated by laughing owls (Sceloglaux albifacies) on the Gouland Downs, Northwest Nelson, South Island. Notornis 48: 225-233.

72. Worthy, T.H., 2001. A new species of Platymantis (Anura: Ranidae) from Quaternary deposits on Viti Levu, Fiji. Palaeontology 44(4): 665-680.

71. Worthy, T.H. 2001. Moa bones from the Matokitoki Gravel Member, Matokitoki Valley Road (Hitchen’s) Quarry, Gisborne. Appendix 1, pp 343-345, In Coleman, J.L., Shane, P.A.R., Grant-Mackie, J.A., Gregory M.R. Identification of a distal ignimbrite in Quaternary strata of the Gisborne area: implications for ignimbrite dispersal, stratigraphy, and paleofauna. New Zealand Journal of Geology and Geophysics 44: 337-345.

70. Holdaway, R.N., T.H. Worthy, A.J.D. Tennyson, 2001. A working list of breeding bird species of the New Zealand region at first human contact. New Zealand journal of zoology 28(2): 119-187.

69. Holdaway, R.N. & T.H. Worthy, 2001. Publications of supplementary data for 14C AMS ages [Forum]. Journal of the Royal Society of New Zealand 31(2): 453-456.

68. Worthy, T.H., 2000. Two late-Glacial avifaunas from eastern North Island, New Zealand – Te Aute Swamp and Wheturau Quarry. Journal of the Royal Society of New Zealand 30(1): 1-26.

67. Worthy, T.H., 2000. New Zealand Pigeon (Hemiphaga novaeseelandiae) on Raoul Island, Kermadec Group. Notornis 47: 36-38.

66. Worthy, T.H., 2000. The fossil megapodes (Aves: Megapodiidae) of Fiji with descriptions of a new genus and two new species.Journal of the Royal Society of New Zealand 30(4): 337-364.

65. Worthy, T.H. & R.N. Holdaway, 2000. Terrestrial Fossil vertebrate faunas from inland Hawke’s Bay, North Island New Zealand. Part 1. Records of the Canterbury Museum 14: 89-154.

64. Steadman, D.W., T.H. Worthy, A.J. Anderson & R. Walter, 2000. New species and records of birds from prehistoric sites on Niue, Southwest Pacific. Wilson Bulletin 112(2): 165-186.

63. Anderson, A., G. Clark & T.H. Worthy, 2000. An inland lapita site in Fiji. Journal of the Polynesian Society 109: 311-316.

62. Worthy, T.H., 1999. What was on the menu – avian extinction in New Zealand. NZ Journal of Archaeology 19: 125-160.

61. Worthy, T.H., A.J. Anderson & R.E. Molnar, 1999. Megafaunal expression in a land without mammals – the first fossil faunas from terrestrial deposits in Fiji. Senckenbergiana biologica 79(2): 237-242.

60. Worthy T.H. & R.N. Holdaway, 1999. Introduction to ‘A preliminary report on the nesting habits of moas on the East Coast of the North Island’ by W. H. Hartree. Notornis 46: 457-460.

59. Worthy, T.H., 1998. Quaternary fossil faunas of Otago, South Island, New Zealand. Journal of the Royal Society of New Zealand 28(3): 421-521.

58. Worthy, T.H., 1998. The Quaternary fossil avifauna of Southland, South Island, New Zealand. Journal of the Royal Society of New Zealand 28(4): 537-589.

57. Worthy, T.H., R. Walter & A.J. Anderson, 1998. Fossil and archaeological avifauna of Niue. Notornis 45: 177-190.

56. Worthy, T.H., 1998. A remarkable fossil and archaeological avifauna from Marfells Beach, Lake Grassmere, South Island, New Zealand. Records of the Canterbury Museum 12(1): 79-176.

55. Worthy, T.H., 1998. Fossils indicate Pelecanoides georgicus had large colonies at Mason Bay, Stewart Island, New Zealand. Notornis 45: 229-246.

54. Worthy, T.H., 1998. Fossil avifaunas from Old Neck and Native Island, Stewart Island – Polynesian middens or natural sites? Records of the Canterbury Museum 12(2): 49-82.

53. Worthy, T.H., 1997. The Quaternary fossil fauna of South Canterbury, South Island, New Zealand. Journal of the Royal Society of New Zealand. 27(1): 67-162.

52. Worthy, T.H., 1997. A survey of modern Laughing Owl Sceloglaux albifacies (Gray, 1844) specimens in collections. Notornis 44: 241-252.

51. Worthy, T.H., 1997. Fossil deposits in the Hodges Creek Cave System, on the northern foothills of Mt Arthur, Nelson, South Island, New Zealand. Notornis 44:111-124.

50. Worthy, T.H., 1997. A mid-Pleistocene rail from New Zealand. Alcheringa 21: 71-78.

49. Worthy, T.H., 1997. The identification of fossil Eudyptes and Megadyptes bones at Marfells Beach, Marlborough, South Island. NZ Natural Sciences 23: 71-85.

48. Holdaway, R.N. & T.H. Worthy, 1997. A reappraisal of the Late Quaternary fossil vertebrates of Pyramid Valley Swamp, North Canterbury, New Zealand. New Zealand Journal of Zoology 24: 69-121.

47. Worthy, T.H., R.N. Holdaway, M.D. Sorenson & A.C. Cooper, 1997. Description of the first complete skeleton of the extinct New Zealand goose Cnemiornis calcitrans Owen, (Aves: Anatidae), and a reassessment of the relationships of Cnemiornis. Journal of Zoology, London 243: 695-723.

46. Worthy, T.H., 1996. Holocene populations of shags Leucocarbo spp. in the Far North, New Zealand. New Zealand Journal of Zoology 23: 89-95.

45. Worthy, T.H., M.J. Daniel & J.E. Hill, 1996. An analysis of skeletal size variation in Mystacina robusta Dwyer, 1962 (Chiroptera: Mystacinidae). New Zealand Journal of Zoology 23: 99-110.

44. Holdaway, R.N. & T.H. Worthy, 1996. Diet and biology of the laughing owl Sceloglaux albifacies (Aves: Strigidae) on Takaka Hill, Nelson, New Zealand. Journal of Zoology London 239: 545-572.

43. Kuschel, G. & T.H. Worthy, 1996. Past distribution of large weevils (Coleoptera: Curculionidae) in the South Island, New Zealand, based on Holocene fossil remains. New Zealand Entomologist 19: 15-22.

42. Worthy, T.H. & R.N. Holdaway, 1996. Quaternary fossil faunas, overlapping taphonomies, and palaeofaunal reconstruction in North Canterbury, South Island, New Zealand. Journal of the Royal Society of New Zealand 26(3): 275-361.

41. Worthy, T.H. & R.N. Holdaway, 1996. Taphonomy of two Holocene microvertebrate deposits, Takaka Hill, Nelson, New Zealand, and identification of the avian predator responsible. Historical Biology 12: 1-24.

40. Worthy, T.H., 1995. Description of some post-cranial bones of Malacorhynchus scarletti, a large extinct Pink-eared duck from New Zealand. Emu 95: 13-22.

39. Worthy, T.H. & R.N. Holdaway, 1995. Quaternary fossil faunas from caves on Mt Cookson, North Canterbury, South Island, New Zealand. Journal of the Royal Society of New Zealand 25(3): 333-370.

38. Vickers-Rich, P., P. Trusler, M.J. Rowley, A, Cooper, G.K. Chambers, W.J. Bock, P.R. Millener, T.H. Worthy & J.C. Yaldwyn, 1995. Morphology, myology, collagen and DNA of a mummified Upland moa, Megalapteryx didinus (Aves: Dinornithiformes) from New Zealand. Tuhinga: Records of the Museum of New Zealand Te Papa Tongarewa 4: 1-26.

37. Worthy, T.H., 1994. Reappraisal of Dinornis (Aves: Dinornithiformes) species – a morphometric analysis. New Zealand Journal of Zoology 21: 113-134.

36. Worthy, T.H., 1994. Holocene breeding of sea lions (Phocarctos hookeri) at Delaware Bay, Nelson, South Island, New Zealand. New Zealand Natural Sciences 21: 61-70.

35. Worthy, T.H., 1994. Late Quaternary changes in the moa fauna on the West Coast of the South Island, New Zealand. Records of the South Australian Museum 27(2): 125-134.

34. Worthy, T.H. & R.N. Holdaway, 1994. Scraps from an owl’s table – predator activity as a significant taphonomic process newly recognised from New Zealand Quaternary deposits. Alcheringa 18: 229-245.

33. Worthy, T.H. & R.N. Holdaway, 1994. Quaternary fossil faunas from caves in Takaka Valley and on Takaka Hill, northwest Nelson, South Island, New Zealand. Journal of the Royal Society of New Zealand 24(3): 297-391.

32. Holdaway, R.N. & Worthy, T.H., 1994. A new fossil species of shearwater Puffinus from the late Quaternary of the South Island, New Zealand, and notes on the biogeography and evolution of the Puffinus gavia superspecies. Emu 94: 201-215.

31. Worthy, T.H., 1993. A review of fossil bird bones from loess deposits in eastern South Island, New Zealand. Records of the Canterbury Museum 10(8): 95-106.

30. Worthy, T.H., 1993. Submarine fossil moa bones from the northern South Island. Journal of the Royal Society of New Zealand 23(3): 255 – 256.

29. Holdaway, R.N. & T.H. Worthy, 1993. First North Island fossil record of kea, and morphological and morphometric comparison of kea and kaka. Notornis 40(2): 95-108.

28. Worthy, T.H. & R.N. Holdaway, 1993. Quaternary fossil faunas from caves in the Punakaiki area, West Coast, South Island, New Zealand. Journal of the Royal Society of New Zealand 23(3): 147 – 254.

27. Cooper, A., I.A.E. Atkinson, W.G. Lee & T.H. Worthy, 1993. Evolution of the moa and their effect on the New Zealand flora. Trends in Ecology and Evolution 8(12): 433 – 442.

26. Worthy, T.H., 1992. A re-examination of the species Euryapteryx geranoides (Owen) including comparisons with other emeiin moas (Aves: Dinornithiformes). Journal of the Royal Society of New Zealand 22(1): 19-40.

25. Worthy, T.H., 1992. Fossil bones of Hooker’s sea lions in New Zealand caves. New Zealand Natural Sciences 19: 31-39.

24. Worthy, T.H., 1991. Fossil skink bones from Northland, New Zealand, including a description of a new Cyclodina, Scincidae. Journal of the Royal Society of New Zealand 21(4): 329 – 348.

23. Worthy, T.H., A.R. Edwards, & P.R. Millener, 1991. The fossil record of moas (Aves: Dinornithiformes) older than the Otira (last) Glaciation. Journal of the Royal Society of New Zealand 21(2): 101-118.

22. Millener, P.R. & T.H. Worthy, 1991. Contributions to New Zealand’s late Quaternary avifauna. II: Dendroscansor decurvirostris, a new genus and species of wren (Aves: Acanthisittidae). Journal of the Royal Society of New Zealand 21(2): 179-200.

21. Worthy, T.H., 1990. An analysis of the distribution and relative abundance of moa species (Aves: Dinornithiformes). New Zealand Journal of Zoology 17: 213-241.

20. Worthy, T.H., 1989. Validation of Pachyornis australis Oliver (Aves: Dinornithiformes), a medium sized moa from the South Island, New Zealand. New Zealand Journal of Geology and Geophysics 32: 255-266.

19. Worthy, T.H., 1989. Mummified moa remains from Mt Owen, northwest Nelson. Notornis 36: 36-38.

18. Worthy, T.H., 1989. Number of presacral vertebrae in Dinornis. Notornis 36: 170.

17. Worthy, T.H., 1989. Moas of the subalpine zone. Notornis 36: 191-196.

16. Worthy, T.H., 1989. The glossohyal and thyroid bone of Aptornis otidiformis. Notornis 36: 248.

15. Worthy, T.H., 1989. Aspects of the biology of two moa species (Aves: Dinornithiformes). New Zealand Journal of Archaeology 11: 77-86.

14. Worthy, T.H., 1989. An analysis of moa bones (Aves: Dinornithiformes) from three lowland North Island swamp sites: Makirikiri, Riverlands and Takapau Road. Journal of the Royal Society of New Zealand 19: 419-432.

13. Worthy, T.H. & D.C. Mildenhall, 1989. A late Otiran-Holocene paleoenvironmental reconstruction based on cave excavations in northwest Nelson, New Zealand. New Zealand Journal of Geology and Geophysics 32: 243-253.

12. Worthy, T.H., 1988. A re-examination of the moa genus Megalapteryx. Notornis 35: 99-108.

11. Worthy, T.H., 1988. An illustrated key to the main leg bones of moas (Aves: Dinornithiformes). National Museum of New Zealand Misc. Publ. Series No. 17 37pp.

10. Worthy, T.H., 1988. Loss of flight ability in the extinct New Zealand duck Euryanas finschi. Journal of Zoology, London 215: 619 – 628.

9. Beauchamp, A. & T.H. Worthy, 1988. Decline in distribution of the takahe Porphyrio (Notornis) mantelli – a re-examination. Journal of the Royal Society of New Zealand 18: 103-112.

8. Cassels, R.J.S., K.L. Jones, A. Walton & T.H. Worthy, 1988. Late prehistoric subsistence practices at Parewanui, lower Rangitikei River, New Zealand. New Zealand Journal of Archaeology 10: 109-128.

7. Worthy, T.H., 1987. Subfossil snipe. Letter, Notornis 34: 343.

6. Worthy, T.H., 1987. Sexual dimorphism and temporal variation in the North Island moa species Euryapteryx curtus (Owen) and Pachyornis mappini Archey. National Museum of New Zealand Records 3(6): 59-70.

5. Worthy, T.H., 1987. Palaeoecological information concerning members of the frog genus Leiopelma: Leiopelmatidae in New Zealand. Journal of the Royal Society of New Zealand 17: 409-420.

4. Worthy, T.H., 1987. Osteological observations of the larger species of skink Cyclodina and the subfossil occurrence of these and the gecko Hoplodactylus duvaucelii in the North Island, New Zealand. New Zealand Journal of Zoology 14: 219-229.

3. Worthy, T.H., 1987. Osteology of Leiopelma (Amphibia: Leiopelmatidae) and descriptions of three new subfossil (Leiopelma) species. Journal of the Royal Society of New Zealand 17: 201-251.

2. Worthy, T.H., 1984. Faunal and floral remains from F1, a cave near Waitomo. Journal of the Royal Society of New Zealand: 14(4): 367-377.

1. Worthy, T.H., 1983. Subfossil insects a key to past diversity. The Weta 6: 42-43.

Peer-reviewed Papers in Conference Proceedings (8)
1. Worthy, T.H., A.J.D. Tennyson, M. Archer & R.P. Scofield 2010. First record of Palaelodus (Aves: Phoenicopteriformes) from New Zealand. In Proceedings of the VII International Meeting of the Society of Avian Paleontology and Evolution, ed. W.E. Boles and T.H. Worthy. Records of the Australian Museum 62(1): 77-88. doi:10.3853/j.0067-1975.62.2010.1545. ERA10C.

2. Scofield, R.P., T.H. Worthy, & A.J.D. Tennyson, 2010. A heron (Aves: Ardeidae) from the Early Miocene St Bathans fauna of southern New Zealand. In Proceedings of the VII International Meeting of the Society of Avian Paleontology and Evolution, ed. W.E. Boles and T.H. Worthy. Records of the Australian Museum 62(1): 89-104. doi:10.3853/j.0067-1975.62.2010.1542. ERA10C.

3. Tennyson, A.J.D., T.H. Worthy, C.M. Jones, R.P. Scofield & S.J. Hand, 2010. Moa’s Ark: Miocene fossils reveal the great antiquity of moa (Aves: Dinornithiformes) in Zealandia. In Proceedings of the VII International Meeting of the Society of Avian Paleontology and Evolution, ed. W.E. Boles and T.H. Worthy. Records of the Australian Museum 62(1): 105-114. doi:10.3853/j.0067-1975.62.2010.1546. ERA10C.

4. Worthy, T.H. & J.X. Zhao, 2006. A late Pleistocene predator-accumulated avifauna from Kids Cave, West Coast, South Island, New Zealand. Alcheringa Special Issue 1: 389-408.

5. Worthy, T.H., M. Bunce, A. Cooper & P. Scofield, 2005. Dinornis – an insular oddity, a taxonomic conundrum reviewed. Alcover, J. A. & Bover, P. (eds), Proceedings of the International Symposium “Insular Vertebrate Evolution: The Palaeontological approach” Monographies de la Societat d’Historia Natural de les Balears 12: 337-390.

6. Worthy, T.H., 1999. The role of climate change versus human impacts – Avian extinction on South Island, New Zealand. Proceedings of the 4th International meeting of the Society of Avian Paleontology and Evolution. Washington, USA, June 1996. Smithsonian Contributions to Paleobiology 89: 111-123.

7. Worthy, T.H. & P. Jouventin, 1999. The fossil avifauna of Amsterdam Island. Proceedings of the 4th International meeting of the Society of Avian Paleontology and Evolution. Washington, USA, June 1996. Smithsonian Contributions to Paleobiology 89: 39-65.

8. Worthy, T.H., 1991. An overview of the taxonomy, fossil history, biology and extinction of moas. In, Bell, B.D., R.O. Cossee, J.E.C. Flux, B.D. Heather, R.A. Hitchmough, C.J.R. Robertson, and M.J. Williams (eds), Acta XX Congressus Internationalis Ornithologici. Vol. 1, pp 555-562.
Books (5)
1. Gill, B.J., B.D. Bell, G.K. Chambers, D.G. Medway, R.L. Palma, R.P. Scofield, A.J.D. Tennyson & T.H. Worthy, 2010. Checklist of the Birds of New Zealand, Norfolk and Macquarie Islands, and the Ross Dependency, Antarctica. 4th Edition. Ornithological Society of New Zealand & Te Papa Press, Wellington, 500 pp.

2. Worthy, T.H., 2008. Tertiary fossil waterfowl (Aves: Anseriformes) of Australia and New Zealand. Thesis for Doctor of Philosophy, University of Adelaide, Adelaide, South Australia; ix + 415 pp. http://digital.library.adelaide.edu.au/dspace/bitstream/2440/50511/2/01front.pdf

3. Worthy, T.H. & R.N. Holdaway, 2002 (9 August). The lost world of the moa: Prehistoric life of New Zealand. Indiana University Press, Indiana. Xxxiii + 718 pp.

4. Worthy, T.H., 1993. Fossils of Honeycomb Hill. Museum of New Zealand Te Papa Tongarewa. 56 pp.

5. Worthy, T.H., 1983. Moas and other subfossils. Waitomo Museum handbook.

Book Chapters (17)
1. Worthy, T.H., 2010. Sections: Dinornithiformes (Pp 11-18), Casuariiformes (Pp 19-23), Anseriformes (pp 30-48), Podicipediformes (Pp 49-51), Accipitriformes (Pp 169-173), Falconiformes (Pp 174-176), Gruiformes (Pp 177-190), Columbiformes (Pp 244-248), Strigiformes (Pp 264-268), Aegothelidae (Pp 269-270), Appendix 1 – Fossil Birds of New Zealand (Pp 323-333). In: Checklist Committee (O.S.N.Z.). 2010. Checklist of the Birds of New Zealand, Norfolk and Macquarie Islands, and the Ross Dependency, Antarctica (4th ed.). Ornithological Society of New Zealand & Te Papa Press, Wellington.

2. Worthy, T.H. & A.J. Anderson, 2009. Palaeofaunal sites and excavations. Pp 19-40, Chapter 2 In: Eds, G Clark and A Anderson, The Early History of Fiji, Terra Australis Volume 31, ANU, The Australian National University E Press, Canberra.

3. Worthy, T.H. & A.J. Anderson, 2009. Results of Palaeofaunal Research. Pp 41-62. Chapter 3, In: Eds, G Clark and A Anderson, The Early History of Fiji, Terra Australis Volume 31, ANU, The Australian National University E Press, Canberra.

4. Worthy, T.H. & G. Clark, 2009. Bird, mammal and reptile remains. Pp 231-258, Chapter 10, In: Eds, G Clark and A Anderson, The Early History of Fiji, Terra Australis Volume 31, ANU, The Australian National University E Press, Canberra.

5. Worthy, T.H., 2009. Fossil Birds, pp 318-326 in R.G. Gillespie & D.A. Clague, eds, Encyclopedia of Islands. University of California Press, Berkley, Los Angeles, London, 1074 pp.

6. King, C.M., C.D. Roberts, B.D. Bell, R.E. Fordyce, R.S. Nicoll, T.H. Worthy, C.D. Paulin, R.A. Hitchmough, I.W. Keyes, A.N. Baker, A.L. Stewart, N. Hiller, B.M. McDowall, R.N. Holdaway, R.P. McPhee, W.W. Schwarzhans, A.J.D. Tennyson, S. Rust & I. McCadie, 2009. Phylum Chordata: Lancelets, Fishes, Amphibians, Reptiles, Birds, and Mammals. New Zealand Inventory of Biodiversity: Volume One: Kingdom Animalia. Ed. D. P. Gordon, Canterbury University Press. Pp 431-527. 566 pp.

7. Holdaway, R.N. & T.H. Worthy, 2008. The Late Quaternary avifauna. Pp, 445-492, in M.J. Winterbourn, G.A. Knox, C.J. Burrows & I.D Marsden (eds), The natural history of Canterbury, 3rd edition. Canterbury University Press and Manaaki Whenua Press, Christchurch, 926 pp.

8. Worthy, T.H. & G.M. Wragg, 2008. A new genus and species of pigeon (Aves: Columbidae) from Henderson Island, Pitcairn Group. Pp 499-510, in: G. Clark, F. Leach, S. O’Connor, (eds), Terra Australis 29. Islands of Inquiry: Colonisation, seafaring and the archaeology of maritime landscapes. ANU E Press, The Australian National University, Canberra. 510 pp.

9. Holdaway, R.N. & T.H. Worthy, 2006. Evolution of New Zealand and its vertebrates. Chapter 6 In: J. Merrick, M. Archer, G.M. Hickey & M.S.Y. Lee, (eds). Evolution and Biogeography of Australasian Vertebrates, Auscipub Pty Ltd: Sydney; pp 117-134.

10. Worthy, T.H. & A.J.D. Tennyson, 2004. Avifaunal assemblages from the Nenega-iti and Onemea sites. Chapter 6. In P.V. Kirch & E. Conte (eds) Archaeological investigations in the Mangareva Islands (Gambier Archipelago), French Polynesia. Berkeley, California, Archaeological Research Facility, University of California; pp. 122-127.

11. Worthy, T., 2004. Moa. In: S. Krech, J.R. McNeill, & C. Merchant (Eds.), Encyclopedia of World Environmental History, 2 vols, Vol 2: 856-858. Routledge, New York/London.

12. Irwin, G.J., R.K. Nichol, M.A. Taylor, T.H. Worthy & I.W.G. Smith, 2004. Faunal remains from Kohika. Chapter 12, pp 198-216. In: Kohika. The archaeology of a late Maori lake village in the Ngati Awa rohe, Bay of Plenty, New Zealand. Ed. G.J. Irwin. Auckland University Press, University of Auckland, Auckland, New Zealand. 266 pp.

13. McGlone, M., P. Wardle, & T.H. Worthy, 2003. Environmental change since the Last Glaciation, Chapter 6, pp105-128, In: The Natural History of Southern New Zealand. Eds J. Darby, R.E. Fordyce, A. Mark, K. Probert & C. Townsend, Otago University Press, Dunedin, New Zealand.

14. Anderson, A., S. Haberle, G. Rojas, A. Seelenfreund, I. Smith, & T. Worthy, 2002. An archaeological exploration of Robinson Crusoe Island, Juan Fernandez Archipelago, Chile. In Fifty years in the field. Essays in honour and celebration of Richard Shutler. JR’s Archaeological career. Eds: S. Bedford, C. Sand & D. Burley, pp 239-249.

15. Anderson, A., L. Ayliffe, D. Questiaux, T. Sorovi-Vunidilo, T.[H.] Worthy, 2001. The terminal age of the Fijian megafauna. Pp 251-264. In: Eds. A. Anderson, I. Lilley, & S. O’Connor, Histories of old ages: Essays in honour of Rhys Jones. Pandanus Books, Research School of Pacific and Asian Studies, The Australian National University.

16. Trewick, S.A. & T.H. Worthy, 2001. Origins and prehistoric ecology of takahe based on morphometric, molecular, and fossil data. Pp 31-48, In: The Takahe: Fifty Years of Conservation Management and Research. Eds: William G. Lee and Ian G. Jamieson. University of Otago Press, Dunedin.

17. Anderson, A., T. Worthy & R., McGovern-Wilson, 1996. Moa remains and taphonomy. Chapter 14, pp 200-213, In. Shag River Mouth, the archaeology of an early Maori village. Anderson, A., Allingham, B. & Smith, I. (eds). Research Papers in Archaeology and Natural History, No. 27, ANH publications, RSPAS, The Australian National University, Canberra, Australia.